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Thus, Activin-A drives pathogenic Th17 cell differentiation, implicating the Activin-A-ALK4-ERK axis as a therapeutic target for Th17 cell-related diseases. To determine the optimal combination of imaging and biochemical biomarkers to predict knee osteoarthritis (OA) progression. Nested case-control study from the FNIH OA Biomarkers Consortium of participants with Kellgren-Lawrence grade 1-3 and complete biomarker data (n=539 to 55. Cases were knees with radiographic and pain progression between 24-48 months from baseline. Radiog