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We recovered four distinct phylogenetic lineages that, despite the overlap of morphometric traits, can be separated phenotypically by subtle but discrete morphological characters. One lineage corresponds to Macrobiotus pallarii, whereas the other three are newly described as Macrobiotus margoae Stec, Vecchi Bartels, sp. nov. from the USA, Macrobiotus ripperi Stec, Vecchi Michalczyk, sp. nov. from Poland and Finland, and Macrobiotus pseudopallarii Stec, Vecchi Michalczyk, sp. nov. from Montenegro. To facilitate species
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