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In this research we tested the role of the MD in timing, utilizing an operant temporal production task in male mice. In this task, the expected timing of offered incentives is indicated by lever pressing. Inactivation of this MD with muscimol produced rightward shifts in peak pressing on probe trials along with increases in peak scatter, thus notably changing both temporal reliability and precision. Optogenetic inhibition of glutamatergic projection neurons in the MD additionally resulted in similar alter