Centrioles are essential organelles that mediate the formation of cilia and flagella, and are vital for many cells during embryonic development. The molecular mechanisms underlying their assembly and function are still poorly understood. Recent evidence suggests that centrioles have a two step biogenesis in most eukaryotes. In the first step a cartwheel consisting of nine microtubule doublets is nucleated at one site to assemble a wall structure in the center. The resulting centriole 50e0806aeb ogyquin
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